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- March 31, 2016.
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- 1Institute of Evolutionary Biology, University of Edinburgh, Kings Buildings, EH9 3JT, United Kingdom
- 2Present Address: Department of Genetics, University of Cambridge, Downing Street, Cambridge, CB2 3EH
- 3Present Address: Life Sciences, University of Sussex, United Kingdom
- 4Department of Biosciences, Centre of Excellence in Biological Interactions, University of Helsinki, Helsinki, Finland
- 5Centre for Immunity, Infection and Evolution, University of Edinburgh, Kings Buildings, EH9 3JT, United Kingdom
- ↵*Author for correspondence: sam.lewis{at}gen.cam.ac.uk