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- June 13, 2017.
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- 1Departamento de Ciencias de la Vida, Universidad de Alcalá, Edificio de Ciencias, Ctra. Madrid-Barcelona km. 33,6, 28871 Alcalá de Henares, Madrid, Spain.
- 2Departamento de Biogeografía y Cambio Global, Museo Nacional de Ciencias Naturales (MNCN-CSIC), C/José Gutiérrez Abascal 2, 28006 Madrid, Spain.
- 3Institute of Plant Sciences, University of Bern, Altenbergrain 21, Bern 3013, Switzerland
- 4Departamento de Biodiversidad y Conservación. Real Jardín Botánico de Madrid (CSIC). 28014 Madrid, España