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- October 31, 2014.
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- Version 1 (April 7, 2014 - 21:05).
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The copyright holder for this preprint is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under a CC-BY 4.0 International license.
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- 1Department of Biological Sciences & Institute for Bioinformatics and Evolutionary Studies, University of Idaho, Moscow, ID 83844 U.S.A.
- 2Department of Biological Sciences, Macquarie University,Sydney, NSW 2109, Australia; rich.fitzjohn{at}gmail.com
- 3School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, NSW 2052 Australia w.cornwell{at}unsw.edu.au
- ↵*Email for correspondence: mwpennell{at}gmail.com