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- June 25, 2015.
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- †Department of Organismic and Evolutionary Biology, Harvard University, Cambridge MA, 02138
- §Max-Planck Odense Center on the Biodemography of Aging, Institute of Biology, University of Southern Denmark, DK-5230 Odense M, Denmark
- ‡Harvard Forest, Harvard University, Petersham MA, 01366
- Corresponding Author: Kolea Zimmerman Address: 16 Divinity Ave, Biolabs 2112, Cambridge MA 02138 Email: kzimmerman{at}fas.harvard.edu