3The Walter and Eliza Hall Institute of Medical Research, 1G Royal Parade, Parkville 3052, Victoria, Australia
4Department of Medical Biology, The University of Melbourne, Parkville 3010, Victoria, Australia
5Department of Gastroenterology, The Royal Melbourne Hospital, Grattan St., Parkville 3050, Victoria, Australia
6Murdoch Children’s Research Institute, Flemington Road, Parkville, Victoria 3050, Australia