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- January 15, 2016.
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- 1Institute of Healthy Ageing, Department of Genetics, Evolution and Environment, University College London, Gower Street, London WC1E 6BT, UK.
- 2Institute of Animal Genetics and Breeding, Sichuan Agricultural University, Chengdu, Sichuan, China.
- 3Berlin Institute of Health, Luisenstraße 56, 10117 Berlin, Germany.
- ↵†For correspondence: yangmingyao{at}sicau.edu.cn, m.piper{at}ucl.ac.uk