1Department of Genetics, Evolution and Environment, University College London, London WC1E 6BT, United Kingdom
4Department of Computer Science, University College London, London WC1E 6BT, United Kingdom
5Department of Ecology and Evolution & Center for Integrative Genomics, University of Lausanne, Biophore, 1015 Lausanne, Switzerland
6Swiss Institute of Bioinformatics, Biophore, 1015 Lausanne, Switzerland
