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- November 3, 2016.
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- 1Department of Biology Pennsylvania State University 408 Life Sciences Building University Park PA 16802-5301
- 3Nicholas School of the Environment Duke University 450 Research Drive Durham, NC 27708-0001
- 4Department of Statistics Pennsylvania State University 411 Thomas Building University Park PA 16802-5301
- ↵2Correspondence: jrl35{at}psu.edu, Phone: (814) 863-5318, Fax: (814) 865-9131