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- December 15, 2016.
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The copyright holder for this preprint is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under a CC-BY-NC-ND 4.0 International license.
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- ↵*Author Correspondence: R. Nakadai, Center for Ecological Research, Kyoto University, Hirano 2-509-3, Otsu, Shiga, 520-2113 Japan, E-mail: r.nakadai@ecology.kyoto-u.ac.jp, Phone: +81-77-549-8019; Fax: +81-77-549-8201