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History
- April 17, 2017.
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The copyright holder for this preprint is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. All rights reserved. No reuse allowed without permission.
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- ↵*To whom correspondence should be addressed. E-mail: othmer{at}umn.edu
The authors declare no conflict of interest.
Authors are listed alphabetically. Jia Gou and Lin Lin contributed equally to this work.