*Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA, 02138, USA
†The Max Planck-Harvard Research Center for the Archaeoscience of the Ancient Mediterranean, Cambridge, MA, 02138, USA and Jena, D-07745, Germany
‡Department of Genetics, Harvard Medical School, Boston, MA, 02115, USA
§Department of Human Evolutionary Biology, Harvard University, Cambridge, MA, 02138, USA
