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- October 17, 2020.
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- 1Department of Integrative Biology, Michigan State University, East Lansing, Michigan
- 2Program in Ecology, Evolutionary Biology, and Behavior, Michigan State University, East Lansing, Michigan
- 3BEACON Center for the Study of Evolution in Action, Michigan State University, East Lansing, Michigan
- 4Mara Hyena Project, Michigan State University, Masai Mara National Reserve, Kenya
- 5School of Biological Sciences, University of Nebraska-Lincoln, Lincoln, Nebraska
- ↵*Corresponding author; email: straussed{at}gmail.com