aCalifornia Institute for Quantitative Biosciences (QB3), University of California, Berkeley, California 94720, USA
bMolecular Biophysics and Integrative Bio-Imaging Division, Lawrence Berkeley National Laboratory, Berkeley, California 94720, USA
dDepartment of Molecular and Cell Biology, University of California, Berkeley, California 94720, USA
fHoward Hughes Medical Institute, University of California, Berkeley, California 94720, USA
