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- September 23, 2020.
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- 1Institute of Biotechnology, University of Helsinki, 00014 Helsinki, Finland
- 2Genomics, Bioinformatics and Evolution. Departament de Genètica i Microbiologia, Universitat Autònoma de Barcelona, 08193, Cerdanyola del Vallès, Spain
- 3Centre de Rercerca Matemàtica, 08193, Cerdanyola del Vallès, Spain
- 4Institute of Molecular and Cell Biology, University of Tartu, 51010 Tartu, Estonia
- ↵*Corresponding authors: d.toddie-moore{at}ed.ac.uk and osamu.shimmi{at}helsinki.fi