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- September 30, 2020.
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- 1Department of Ecology and Evolution, University of Chicago, Chicago, IL 60637, USA
- 2Center for the Physics of Evolving Systems, University of Chicago, Chicago, IL 60637, USA
- 3Department of Physics, University of Illinois at Urbana-Champaign, Urbana, IL 61801, USA
- 4Department of Engineering Sciences and Applied Mathematics, Northwestern University, Evanston, IL 60208, USA
- 5Department of Molecular Biosciences, Northwestern University, Evanston, IL 60208, USA
- 6NSF-Simons Center for Quantitative Biology, Northwestern University, Northwestern University, Evanston, IL 60208, USA
- ↵*To whom correspondence should be addressed; E-mail: madhav.mani{at}gmail.com, seppe.kuehn{at}gmail.com.