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- September 4, 2021.
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- Shoji Hayashi1,2,*,**,
- Mugino O. Kubo3,*,**,
- Marcelo R. Sánchez-Villagra4,
- Hiroyuki Taruno5,
- Masako Izawa6,7,
- Tsunehiro Shiroma6,
- Takayoshi Nakano2 and
- Masaki Fujita8
- 1Faculty of Biosphere-Geosphere Science, Okayama University of Science, Kita-ku Ridaicho 1-1, Okayama, 700-0005, Japan
- 2Division of Materials and Manufacturing Science, Graduate School of Engineering, Osaka University, Yamada-Oka, Suita, Osaka, 565-0871, Japan
- 3Department of Natural Environmental Studies, Graduate School of Frontier Sciences, The University of Tokyo, 5-1-5 Kashiwanoha, Kashiwa, Chiba 277-8563, Japan
- 4Paläontologisches Institut und Museum der Universität Zürich, Karl Schmid-Strasse 4, CH-8006 Zürich, Switzerland
- 5Osaka Museum of Natural History, Nagai Park 1-23, Higashi-Sumiyoshi-ku, Osaka, 546-0034, Japan
- 6Faculty of Science, University of the Ryukyus, Okinawa 903-0213, Japan
- 7Kitakyushu Museum of Natural History & Human History, 2-4-1 Higashida, Yahatahigashi-ku, Kitakyushu 805-0071, Japan
- 8Department of Anthropology, National Museum of Nature and Science, 4-1-1 Amakubo, Tsukuba-shi, Ibaraki 305-0005, Japan
- ↵**Authors for correspondence: Shoji Hayashi e-mail: hayashi{at}big.ous.ac.jp, Mugino O. Kubo e-mail: mugino{at}k.u-tokyo.ac.jp
↵* Both authors contributed equally to this work.