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- March 23, 2021.
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- 1Animal Ecology, Department of Biology, Utrecht University, Padualaan 8, 3584 CA Utrecht, The Netherlands
- 2Department for the Ecology of Animal Societies, Max Planck of Animal Behavior, Bücklestraße 5a, 78467, Konstanz, Germany
- 3Behavioral Ecology and Ecophysiology Group, Department of Biology, University of Antwerp, Universiteitsplein 1, 2610 Wilrijk, Belgium
- 4Centre for Research and Conservation, Royal Zoological Society of Antwerp, K. Astridplein 26, B 2018 Antwerp, Belgium
- ↵*Corresponding author: ejcvanleeuwen{at}gmail.com
↵† contributed equally