1Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, Japan
4Department of Integrated Biosciences, Graduate School of Frontier Sciences, The University of Tokyo, Chiba, Japan
5Department of Computational Biology and Medical Sciences, Graduate School of Frontier Sciences, The University of Tokyo, Chiba, Japan
6Atmosphere and Ocean Research Institute, The University of Tokyo, Chiba, Japan
7Institute for Quantitative Biosciences, The University of Tokyo, Tokyo, Japan
8Collaborative Research Institute for Innovative Microbiology, The University of Tokyo, Tokyo, Japan