1Division of Systems Virology, Department of Microbiology and Immunology, The Institute of Medical Science, The University of Tokyo, Tokyo, Japan
3Graduate School of Medicine, The University of Tokyo, Tokyo, Japan
4Graduate School of Frontier Sciences, The University of Tokyo, Kashiwa, Japan
6International Research Center for Infectious Diseases, The Institute of Medical Science, The University of Tokyo, Tokyo, Japan
7International Vaccine Design Center, The Institute of Medical Science, The University of Tokyo, Tokyo, Japan
8Collaboration Unit for Infection, Joint Research Center for Human Retrovirus infection, Kumamoto University, Kumamoto, Japan
9CREST, Japan Science and Technology Agency, Kawaguchi, Japan
