1Department of Developmental, Molecular & Chemical Biology, Tufts University School of Medicine, Boston, MA 02111, USA
2Program of Genetics, Molecular & Cellular Biology, Tufts Graduate School of Biomedical Sciences, Boston, MA 02111, USA
4Laboratory for the Convergence of Biomedical, Physical, and Engineering Sciences, Tufts University School of Medicine, Boston, MA 02111, USA