1Department of Anatomy, University of Otago, Dunedin, New Zealand
2Department of Fish Ecology and Evolution, Eawag, Swiss Federal Institute of Aquatic Science and Technology, Centre of Ecology, Evolution and Biochemistry, Seestrasse 79, 6047 Kastanienbaum, Lucerne, Switzerland
3Department of Ecology, Universität Innsbruck, Technikerstr. 25, 6020 Innsbruck, Austria
