1Department of Molecular Biosciences, Northwestern University, Evanston, IL 60208, USA
2NSF-Simons Center for Quantitative Biology, Northwestern University, Evanston, IL 60208, USA
6Department of Engineering Sciences and Applied Mathematics, Northwestern University, Evanston, IL 60208, USA
7Department of Physics and Astronomy, Northwestern University, Evanston, IL 60208, USA
8Northwestern Institute on Complex Systems, Northwestern University, Evanston, IL 60208, USA
9Santa Fe Institute, Santa Fe, NM 87501, USA
