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The genetic architecture of the human cerebral cortex

Katrina L. Grasby, Neda Jahanshad, Jodie N. Painter, Lucía Colodro-Conde, Janita Bralten, Derrek P. Hibar, Penelope A. Lind, Fabrizio Pizzagalli, Christopher R.K. Ching, Mary Agnes B. McMahon, Natalia Shatokhina, Leo C.P. Zsembik, Ingrid Agartz, Saud Alhusaini, Marcio A.A. Almeida, Dag Alnæs, Inge K. Amlien, Micael Andersson, Tyler Ard, Nicola J. Armstrong, Allison Ashley-Koch, Manon Bernard, Rachel M. Brouwer, Elizabeth E.L. Buimer, Robin Bülow, Christian Bürger, Dara M. Cannon, Mallar Chakravarty, Qiang Chen, Joshua W. Cheung, Baptiste Couvy-Duchesne, Anders M. Dale, Shareefa Dalvie, Tânia K. de Araujo, Greig I. de Zubicaray, Sonja M.C. de Zwarte, Anouk den Braber, Nhat Trung Doan, Katharina Dohm, Stefan Ehrlich, Hannah-Ruth Engelbrecht, Susanne Erk, Chun Chieh Fan, Iryna O. Fedko, Sonya F. Foley, Judith M. Ford, Masaki Fukunaga, Melanie E. Garrett, Tian Ge, Sudheer Giddaluru, Aaron L. Goldman, Nynke A. Groenewold, Dominik Grotegerd, Tiril P. Gurholt, Boris A. Gutman, Narelle K. Hansell, Mathew A. Harris, Marc B. Harrison, Courtney C. Haswell, Michael Hauser, Stefan Herms, Dirk J. Heslenfeld, New Fei Ho, David Hoehn, Per Hoffmann, Laurena Holleran, Martine Hoogman, Jouke-Jan Hottenga, Masashi Ikeda, Deborah Janowitz, Iris E. Jansen, Tianye Jia, Christiane Jockwitz, Ryota Kanai, Sherif Karama, Dalia Kasperaviciute, Tobias Kaufmann, Sinead Kelly, Masataka Kikuchi, Marieke Klein, Michael Knapp, Annchen R. Knodt, Bernd Krämer, Max Lam, Thomas M. Lancaster, Phil H. Lee, Tristram A. Lett, Lindsay B. Lewis, Iscia Lopes-Cendes, Michelle Luciano, Fabio Macciardi, Andre F. Marquand, Samuel R. Mathias, Tracy R. Melzer, Yuri Milaneschi, Nazanin Mirza-Schreiber, Jose C.V. Moreira, Thomas W. Mühleisen, Bertram Müller-Myhsok, Pablo Najt, Soichiro Nakahara, Kwangsik Nho, Loes M. Olde Loohuis, Dimitri Papadopoulos Orfanos, John F. Pearson, Toni L. Pitcher, Benno Pütz, Anjanibhargavi Ragothaman, Faisal M. Rashid, Ronny Redlich, Céline S. Reinbold, Jonathan Repple, Geneviève Richard, Brandalyn C. Riedel, Shannon L. Risacher, Cristiane S. Rocha, Nina Roth Mota, Lauren Salminen, Arvin Saremi, Andrew J. Saykin, Fenja Schlag, Lianne Schmaal, Peter R. Schofield, Rodrigo Secolin, Chin Yang Shapland, Li Shen, Jean Shin, Elena Shumskaya, Ida E. Sønderby, Emma Sprooten, Lachlan T. Strike, Katherine E. Tansey, Alexander Teumer, Anbupalam Thalamuthu, Sophia I. Thomopoulos, Diana Tordesillas-Gutiérrez, Jessica A. Turner, Anne Uhlmann, Costanza Ludovica Vallerga, Dennis van der Meer, Marjolein M.J. van Donkelaar, Liza van Eijk, Theo G.M. van Erp, Neeltje E.M. van Haren, Daan van Rooij, Marie-José van Tol, Jan H. Veldink, Ellen Verhoef, Esther Walton, Mingyuan Wang, Yunpeng Wang, Joanna M. Wardlaw, Wei Wen, Lars T. Westlye, Christopher D. Whelan, Stephanie H. Witt, Katharina Wittfeld, Christiane Wolf, Thomas Wolfers, Clarissa L. Yasuda, Dario Zaremba, Zuo Zhang, Alyssa H. Zhu, Marcel P. Zwiers, Eric Artiges, Amelia A. Assareh, Rosa Ayesa-Arriola, Aysenil Belger, Christine L. Brandt, Gregory G. Brown, Sven Cichon, Joanne E. Curran, Gareth E. Davies, Franziska Degenhardt, Bruno Dietsche, Srdjan Djurovic, Colin P. Doherty, Ryan Espiritu, Daniel Garijo, Yolanda Gil, Penny A. Gowland, Robert C. Green, Alexander N. Häusler, Walter Heindel, Beng-Choon Ho, Wolfgang U. Hoffmann, Florian Holsboer, Georg Homuth, Norbert Hosten, Clifford R. Jack Jr., MiHyun Jang, Andreas Jansen, Knut Kolskår, Sanne Koops, Axel Krug, Kelvin O. Lim, Jurjen J. Luykx, Daniel H. Mathalon, Karen A. Mather, Venkata S. Mattay, Sarah Matthews, Jaqueline Mayoral Van Son, Sarah C. McEwen, Ingrid Melle, Derek W. Morris, Bryon A. Mueller, Matthias Nauck, Jan E. Nordvik, Markus M. Nöthen, Daniel S. O’Leary, Nils Opel, Marie - Laure Paillère Martinot, G. Bruce Pike, Adrian Preda, Erin B. Quinlan, Varun Ratnakar, Simone Reppermund, Vidar M. Steen, Fábio R. Torres, Dick J. Veltman, James T. Voyvodic, Robert Whelan, Tonya White, Hidenaga Yamamori, Oscar L. Lopez, Hieab H.H. Adams, Joshua C. Bis, Stephanie Debette, Charles Decarli, Myriam Fornage, Vilmundur Gudnason, Edith Hofer, M. Arfan Ikram, Lenore Launer, W. T. Longstreth, Bernard Mazoyer, Thomas H. Mosley, Gennady V. Roshchupkin, Claudia L. Satizabal, Reinhold Schmidt, Sudha Seshadri, Qiong Yang, The Alzheimer’s Disease Neuroimaging Initiative, CHARGE Consortium, EPIGEN consortium, IMAGEN consortium, SYS consortium, Parkinson’s Progression Markers Initiative (PPMI), Marina K.M. Alvim, David Ames, Tim J. Anderson, Ole A. Andreassen, Alejandro Arias-Vasquez, Mark E. Bastin, Bernhard T. Baune, John Blangero, Dorret I. Boomsma, Henry Brodaty, Han G. Brunner, Randy L. Buckner, Jan K. Buitelaar, Juan R. Bustillo, Wiepke Cahn, Vince Calhoun, Xavier Caseras, Svenja Caspers, Gianpiero L. Cavalleri, Fernando Cendes, Benedicto Crespo-Facorro, John C. Dalrymple-Alford, Udo Dannlowski, Eco J.C. de Geus, Ian J. Deary, Chantal Depondt, Sylvane Desrivières, Gary Donohoe, Thomas Espeseth, Guillén Fernández, Simon E. Fisher, Herta Flor, Andreas J. Forstner, Clyde Francks, Barbara Franke, David C. Glahn, Randy L. Gollub, Hans J. Grabe, Oliver Gruber, Asta K. Håberg, Ahmad R. Hariri, Catharina A. Hartman, Ryota Hashimoto, Andreas Heinz, Manon H.J. Hillegers, Pieter J. Hoekstra, Avram J. Holmes, L. Elliot Hong, William D. Hopkins, Hilleke E. Hulshoff Pol, Terry L. Jernigan, Erik G. Jönsson, René S. Kahn, Martin A. Kennedy, Tilo T.J. Kircher, Peter Kochunov, John B.J. Kwok, Stephanie Le Hellard, Nicholas G. Martin, Jean-Luc Martinot, Colm McDonald, Katie L. McMahon, Andreas Meyer-Lindenberg, Rajendra A. Morey, Lars Nyberg, Jaap Oosterlaan, Roel A. Ophoff, Tomas Paus, Zdenka Pausova, Brenda W.J.H. Penninx, Tinca J.C. Polderman, Danielle Posthuma, Marcella Rietschel, Joshua L. Roffman, Laura M. Rowland, Perminder S. Sachdev, Philipp G. Sämann, Gunter Schumann, Kang Sim, Sanjay M. Sisodiya, Jordan W. Smoller, Iris E. Sommer, Beate St Pourcain, Dan J. Stein, Arthur W. Toga, Julian N. Trollor, Nic J.A. Van der Wee, Dennis van’t Ent, Henry Völzke, Henrik Walter, Bernd Weber, Daniel R. Weinberger, Margaret J. Wright, Juan Zhou, Jason L. Stein, Paul M. Thompson, Sarah E. Medland, on behalf of the Enhancing NeuroImaging Genetics through Meta-Analysis Consortium - Genetics working group
doi: https://doi.org/10.1101/399402
Katrina L. Grasby
Psychiatric Genetics, QIMR Berghofer Medical Research Institute, Brisbane, Australia
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Neda Jahanshad
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Jodie N. Painter
Psychiatric Genetics, QIMR Berghofer Medical Research Institute, Brisbane, Australia
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Lucía Colodro-Conde
Psychiatric Genetics, QIMR Berghofer Medical Research Institute, Brisbane, Australia
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Janita Bralten
Department of Human Genetics, Radboud university medical center, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The Netherlands
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Derrek P. Hibar
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USANeuroscience Biomarkers, Janssen Research and Development, LLC, San Diego, USA
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Penelope A. Lind
Psychiatric Genetics, QIMR Berghofer Medical Research Institute, Brisbane, Australia
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Fabrizio Pizzagalli
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Christopher R.K. Ching
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USAGraduate Interdepartmental Program in Neuroscience, University of California Los Angeles, Los Angeles, USA
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Mary Agnes B. McMahon
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Natalia Shatokhina
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Leo C.P. Zsembik
Department of Genetics & UNC Neuroscience Center, University of North Carolina at Chapel Hill, Chapel Hill, USA
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Ingrid Agartz
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, NorwayDepartment of Psychiatric Research, Diakonhjemmet Hospital, Oslo, NorwayCentre for Psychiatric Research, Department of Clinical Neuroscience, Karolinska Institutet, Stockholm, Sweden
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Saud Alhusaini
Department of Molecular and Cellular Therapeutics, Royal College of Surgeons in Ireland, Dublin, IrelandNeurology Department, University of California at San Francisco, San Francisco, USA
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Marcio A.A. Almeida
Department of Human Genetics and South Texas Diabetes and Obesity Institute, University of Texas Rio Grande Valley School of Medicine, Brownsville, USA
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Dag Alnæs
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, Norway
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Inge K. Amlien
Centre for Lifespan Changes in Brain and Cognition, Department of Psychology, University of Oslo, Oslo, Norway
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Micael Andersson
Department of Integrative Medical Biology, Umeå University, Umeå, SwedenUmeå Center for Functional Brain Imaging, Umeå University, Umeå, Sweden
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Tyler Ard
Laboratory of Neuro Imaging, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of the University of Southern California, Los Angeles, USA
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Nicola J. Armstrong
Mathematics and Statistics, Murdoch University, Murdoch, Australia
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Allison Ashley-Koch
Duke Molecular Physiology Institute, Duke University Medical Center, Durham, USA
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Manon Bernard
The Hospital for Sick Children, University of Toronto, Toronto, Canada
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Rachel M. Brouwer
Department of Psychiatry, Brain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The Netherlands
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Elizabeth E.L. Buimer
Department of Psychiatry, Brain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The Netherlands
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Robin Bülow
Institute for Radiology and Neuroradiology, University Medicine, Ernst-Moritz-Arndt University, Greifswald, Germany
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Christian Bürger
Department of Psychiatry, University of Münster, Münster, Germany
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Dara M. Cannon
Centre for Neuroimaging & Cognitive Genomics, National University of Ireland Galway, Galway, Ireland
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Mallar Chakravarty
Douglas Mental Health University Institute, McGill University, Montreal, CanadaDepartments of Psychiatry and Biological and Biomedical Engineering, McGill University, Montreal, Canada
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Qiang Chen
Lieber Institute for Brain Development, Baltimore, USA
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Joshua W. Cheung
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Baptiste Couvy-Duchesne
Institute for Molecular Bioscience, The University of Queensland, Brisbane, AustraliaGenetic Epidemiology, QIMR Berghofer Medical Research Institute, Brisbane, AustraliaQueensland Brain Institute, University of Queensland, St Lucia, Australia
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Anders M. Dale
Department of Neurosciences, University of California, San Diego, La Jolla, USADepartment of Radiology, University of California San Diego, San Diego, USA
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Shareefa Dalvie
Department of Psychiatry and Mental Health, University of Cape Town, Cape Town, South Africa
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Tânia K. de Araujo
Department of Medical Genetics, School of Medical Sciences, University of Campinas - UNICAMP, Campinas, Brazil
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Greig I. de Zubicaray
Faculty of Health, Institute of Health and Biomedical Innovation, Queensland University of Technology, Brisbane, Australia
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Sonja M.C. de Zwarte
Department of Psychiatry, Brain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The Netherlands
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Anouk den Braber
Department of Biological Psychology, Vrije Universiteit Amsterdam, Amsterdam, The NetherlandsAlzheimer Center Amsterdam, Department of Neurology, Amsterdam Neuroscience, Vrije Universiteit Amsterdam, Amsterdam UMC, Amsterdam, The Netherlands
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Nhat Trung Doan
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, Norway
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Katharina Dohm
Department of Psychiatry, University of Münster, Münster, Germany
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Stefan Ehrlich
Division of Psychological & Social Medicine and Developmental Neurosciences, Technische Universität Dresden, Dresden, Germany
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Hannah-Ruth Engelbrecht
Division of Human Genetics, Institute of Infectious Disease and Molecular Medicine, University of Cape Town, Cape Town, South Africa
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Susanne Erk
Division of Mind and Brain Research, Department of Psychiatry and Psychotherapy, Campus Charité Mitte, Charité - Universitätsmedizin Berlin corporate member of Freie Universität Berlin, Humboldt-Universität zu Berlin, and Berlin Institute of Health, Berlin, Germany
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Chun Chieh Fan
Department of Cognitive Science, University of California San Diego, San Diego, USA
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Iryna O. Fedko
Department of Biological Psychology, Vrije Universiteit Amsterdam, Amsterdam, The Netherlands
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Sonya F. Foley
Cardiff University Brain Research Imaging Centre, Cardiff University, Cardiff, UK
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Judith M. Ford
San Francisco Veterans Administration Medical Center, San Francisco, USA
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Masaki Fukunaga
Division of Cerebral Integration, National Institute for Physiological Sciences, Okazaki, Japan
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Melanie E. Garrett
Duke Molecular Physiology Institute, Duke University Medical Center, Durham, USA
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Tian Ge
Psychiatric and Neurodevelopmental Genetics Unit, Center for Genomic Medicine, Massachusetts General Hospital, Boston, USAAthinoula A Martinos Center for Biomedical Imaging, Massachusetts General Hospital, Boston, USA
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Sudheer Giddaluru
NORMENT - K.G. Jebsen Centre for Psychosis Research, Department of Clinical Science, University of Bergen, Norway
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Aaron L. Goldman
Lieber Institute for Brain Development, Baltimore, USA
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Nynke A. Groenewold
Department of Psychiatry and Mental Health, University of Cape Town, Cape Town, South Africa
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Dominik Grotegerd
Department of Psychiatry, University of Münster, Münster, Germany
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Tiril P. Gurholt
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, NorwayDepartment of Psychiatric Research, Diakonhjemmet Hospital, Oslo, Norway
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Boris A. Gutman
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USADepartment of Biomedical Engineering, Illinois Institute of Technology, Chicago, USA
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Narelle K. Hansell
Queensland Brain Institute, University of Queensland, St Lucia, Australia
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Mathew A. Harris
Centre for Clinical Brain Sciences and Edinburgh Imaging, University of Edinburgh, Edinburgh, UKDivision of Psychiatry, University of Edinburgh, Edinburgh, UK
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Marc B. Harrison
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Courtney C. Haswell
Duke UNC Brain Imaging and Analysis Center, Duke University Medical Center, Durham, USAMental Illness Research Education and Clinical Center for Post Deployment Mental Health, Durham VA Medical Center, Durham, USA
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Michael Hauser
Duke Molecular Physiology Institute, Duke University Medical Center, Durham, USA
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Stefan Herms
Department of Biomedicine, University of Basel, Basel, SwitzerlandInstitute of Medical Genetics and Pathology, University Hospital Basel, Basel, SwitzerlandDepartment of Genomics, Life & Brain Research Center, University of Bonn, Bonn, Germany
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Dirk J. Heslenfeld
Department of Cognitive and Clinical Neuropsychology, Vrije Universiteit Amsterdam, Amsterdam, The Netherlands
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New Fei Ho
Research Division, Institute of Mental Health, Singapore, Singapore
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David Hoehn
Max Planck Institute of Psychiatry, Munich, Germany
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Per Hoffmann
Department of Biomedicine, University of Basel, Basel, SwitzerlandInstitute of Medical Genetics and Pathology, University Hospital Basel, Basel, SwitzerlandInstitute of Human Genetics, University of Bonn, School of Medicine & University Hospital Bonn, Bonn, Germany
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Laurena Holleran
Centre for Neuroimaging & Cognitive Genomics, National University of Ireland Galway, Galway, Ireland
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Martine Hoogman
Department of Human Genetics, Radboud university medical center, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The Netherlands
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Jouke-Jan Hottenga
Department of Biological Psychology, Vrije Universiteit Amsterdam, Amsterdam, The Netherlands
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Masashi Ikeda
Department of Psychiatry, Fujita Health University School of Medicine, Toyoake, Japan
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Deborah Janowitz
Department of Psychiatry and Psychotherapy, University Medicine Greifswald, Greifswald, Germany
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Iris E. Jansen
Complex Trait Genetics, Center for Neurogenomics and Cognitive Research, Vrije Universiteit Amsterdam, Amsterdam, The NetherlandsDepartment of Neurology, Alzheimer Center, Amsterdam Neuroscience, Vrije Universiteit Medical Center, Vrije Universiteit Amsterdam, Amsterdam, The Netherlands
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Tianye Jia
Institute of Science and Technology for Brain-Inspired Intelligence, Fudan University, Shanghai, ChinaKey Laboratory of Computational Neuroscience and Brain-Inspired Intelligence (Fudan University), Ministry of Education, Shanghai, ChinaCentre for Population Neuroscience and Precision Medicine (PONS), Institute of Psychiatry, Psychology and Neuroscience, King’s College London, London, UK
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Christiane Jockwitz
Institute of Neuroscience and Medicine (INM-1), Research Centre Jülich, Jülich, GermanyDepartment of Psychiatry, Psychotherapy and Psychosomatics, Medical Faculty, RWTH Aachen University, Aachen, GermanyJARA-BRAIN, Jülich-Aachen Research Alliance, Jülich, Germany
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Ryota Kanai
Department of Neuroinformatics, Araya, Inc, Tokyo, JapanSackler Centre for Consciousness Science, School of Psychology, University of Sussex, Falmer, UKEarth-Life Science Institute, Tokyo Institute of Technology, Tokyo, Japan
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Sherif Karama
Department of Psychiatry, McGill University, Montreal, CanadaMcConnell Brain Imaging Center, Montreal Neurological Institute, Montreal, CanadaDouglas Mental Health University Institute, McGill University, Montreal, Canada
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Dalia Kasperaviciute
Department of Clinical and Experimental Epilepsy, UCL Institute of Neurology, London, UK
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Tobias Kaufmann
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, Norway
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Sinead Kelly
Public Psychiatry Division, Massachusetts Mental Health Center, Beth Israel Deaconess Medical Center, Harvard Medical School, Boston, USAPsychiatry Neuroimaging Laboratory, Department of Psychiatry, Brigham and Women’s Hospital, Harvard Medical School, Boston, USA
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Masataka Kikuchi
Department of Genome Informatics, Graduate School of Medicine, Osaka University, Suita, Japan
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Marieke Klein
Department of Human Genetics, Radboud university medical center, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The NetherlandsDepartment of Psychiatry, Brain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The Netherlands
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Michael Knapp
Department of Medical Biometry, Informatics and Epidemiology, University Hospital Bonn, Germany
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Annchen R. Knodt
Department of Psychology and Neuroscience, Duke University, Durham, USA
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Bernd Krämer
Section for Experimental Psychopathology and Neuroimaging, Department of General Psychiatry, Heidelberg University Hospital, Heidelberg, GermanyCentre for Translational Research in Systems Neuroscience and Psychiatry, Department of Psychiatry & Psychotherapy, University Medical Center Göttingen, Göttingen, Germany
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Max Lam
Research Division, Institute of Mental Health, Singapore, SingaporeHuman Genetics, Genome Institute of Singapore, Singapore, Singapore
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Thomas M. Lancaster
Cardiff University Brain Research Imaging Centre, Cardiff University, Cardiff, UKMRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University, Cardiff, UK
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Phil H. Lee
Psychiatric and Neurodevelopmental Genetics Unit, Center for Genomic Medicine, Massachusetts General Hospital, Boston, USADepartment of Psychiatry, Harvard Medical School, Boston, USA
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Tristram A. Lett
Division of Mind and Brain Research, Department of Psychiatry and Psychotherapy, Campus Charité Mitte, Charité - Universitätsmedizin Berlin corporate member of Freie Universität Berlin, Humboldt-Universität zu Berlin, and Berlin Institute of Health, Berlin, Germany
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Lindsay B. Lewis
McGill Centre for Integrative Neuroscience, McGill University, Montreal, CanadaMcConnell Brain Imaging Center, Montreal Neurological Institute, Montreal, Canada
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Iscia Lopes-Cendes
Department of Medical Genetics, School of Medical Sciences, University of Campinas - UNICAMP, Campinas, BrazilBRAINN - Brazilian Institute of Neuroscience and Neurotechnology, Campinas, Brazil
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Michelle Luciano
Department of Psychology, University of Edinburgh, Edinburgh, UKCentre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, UK
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Fabio Macciardi
Department of Psychiatry and Human Behavior, School of Medicine University of California, Irvine, Irvine, USA
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Andre F. Marquand
Department of Cognitive Neuroscience, Radboud university medical center, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The Netherlands
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Samuel R. Mathias
Department of Psychiatry, Yale University School of Medicine, New Haven, USAOlin Neuropsychiatric Research Center, Institute of Living, Hartford Hospital, Hartford, USA
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Tracy R. Melzer
Department of Medicine, University of Otago, Christchurch, Christchurch, New ZealandNew Zealand Brain Research Institute, Christchurch, New ZealandBrain Research New Zealand - Rangahau Roro Aotearoa, Christchurch, New Zealand
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Yuri Milaneschi
Amsterdam UMC, Vrije Universiteit, Psychiatry, Amsterdam Public Health and Amsterdam Neuroscience and GGZinGeest, Amsterdam, The Netherlands
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Nazanin Mirza-Schreiber
Max Planck Institute of Psychiatry, Munich, Germany
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Jose C.V. Moreira
BRAINN - Brazilian Institute of Neuroscience and Neurotechnology, Campinas, BrazilIC - Institute of Computing, Campinas, Brazil
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Thomas W. Mühleisen
Institute of Neuroscience and Medicine (INM-1), Research Centre Jülich, Jülich, GermanyDepartment of Biomedicine, University of Basel, Basel, SwitzerlandCécile and Oskar Vogt Institute of Brain Research, Medical Faculty, Heinrich Heine University, Düsseldorf, Germany
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Bertram Müller-Myhsok
Max Planck Institute of Psychiatry, Munich, GermanyMunich Cluster for Systems Neurology (SyNergy), Munich, GermanyInstitute of Translational Medicine, Liverpool, United Kingdom
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Pablo Najt
Centre for Neuroimaging & Cognitive Genomics, National University of Ireland Galway, Galway, Ireland
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Soichiro Nakahara
Department of Psychiatry and Human Behavior, School of Medicine University of California, Irvine, Irvine, USADrug Discovery Research, Astellas Pharmaceuticals, 21 Miyukigaoka, Tsukuba, Ibaraki 305-8585, Japan
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Kwangsik Nho
Department of Radiology and Imaging Sciences, Indiana University School of Medicine, Indianapolis, USA
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Loes M. Olde Loohuis
Center for Neurobehavioral Genetics, University of California Los Angeles, Los Angeles, USA
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Dimitri Papadopoulos Orfanos
NeuroSpin, CEA, Université Paris-Saclay, Gif-sur-Yvette, France
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John F. Pearson
Biostatistics and Computational Biology Unit, University of Otago Christchurch, Christchurch, New ZealandDepartment of Pathology and Biomedical Science, University of Otago, Christchurch, Christchurch, New Zealand
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Toni L. Pitcher
Department of Medicine, University of Otago, Christchurch, Christchurch, New ZealandNew Zealand Brain Research Institute, Christchurch, New ZealandBrain Research New Zealand - Rangahau Roro Aotearoa, Christchurch, New Zealand
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Benno Pütz
Max Planck Institute of Psychiatry, Munich, Germany
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Anjanibhargavi Ragothaman
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Ronny Redlich
Department of Psychiatry, University of Münster, Münster, Germany
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Department of Biomedicine, University of Basel, Basel, SwitzerlandInstitute of Medical Genetics and Pathology, University Hospital Basel, Basel, Switzerland
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Jonathan Repple
Department of Psychiatry, University of Münster, Münster, Germany
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Geneviève Richard
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, NorwayDepartment of Psychology, University of Oslo, Oslo, NorwaySunnaas Rehabilitation Hospital HT, Nesodden, Norway
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Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USADepartment of Radiology and Imaging Sciences, Indiana University School of Medicine, Indianapolis, USA
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Shannon L. Risacher
Department of Radiology and Imaging Sciences, Indiana University School of Medicine, Indianapolis, USA
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Cristiane S. Rocha
Department of Medical Genetics, School of Medical Sciences, University of Campinas - UNICAMP, Campinas, BrazilBRAINN - Brazilian Institute of Neuroscience and Neurotechnology, Campinas, Brazil
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Nina Roth Mota
Department of Human Genetics, Radboud university medical center, Nijmegen, The NetherlandsDepartment of Psychiatry, Radboud university medical center, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The Netherlands
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Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Andrew J. Saykin
Department of Radiology and Imaging Sciences, Indiana University School of Medicine, Indianapolis, USADepartment of Medical and Molecular Genetics, Indiana University School of Medicine, Indianapolis, USA
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Fenja Schlag
Language and Genetics Department, Max Planck Institute for Psycholinguistics, Nijmegen, The Netherlands
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Lianne Schmaal
Orygen, The National Centre of Excellence for Youth Mental Health, Melbourne, AustraliaThe Centre for Youth Mental Health, University of Melbourne, Melbourne, AustraliaDepartment of Psychiatry, Vrije Universiteit University Medical Center, Vrije Universiteit Amsterdam, Amsterdam, The Netherlands
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Peter R. Schofield
Neuroscience Research Australia, Sydney, AustraliaSchool of Medical Sciences, University of New South Wales, Sydney, Australia
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Rodrigo Secolin
Department of Medical Genetics, School of Medical Sciences, University of Campinas - UNICAMP, Campinas, BrazilBRAINN - Brazilian Institute of Neuroscience and Neurotechnology, Campinas, Brazil
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Chin Yang Shapland
Language and Genetics Department, Max Planck Institute for Psycholinguistics, Nijmegen, The Netherlands
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Li Shen
Department of Biostatistics, Epidemiology and Informatics, University of Pennsylvania, Philadelphia, USA
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Jean Shin
The Hospital for Sick Children, University of Toronto, Toronto, CanadaPopulation Neuroscience & Developmental Neuroimaging, Bloorview Research Institute, University of Toronto, East York, Canada
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Elena Shumskaya
Department of Human Genetics, Radboud university medical center, Nijmegen, The NetherlandsDonders Centre for Cognitive Neuroimaging, Radboud University, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The Netherlands
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Ida E. Sønderby
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, Norway
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Emma Sprooten
Donders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The Netherlands
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Lachlan T. Strike
Queensland Brain Institute, University of Queensland, St Lucia, Australia
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Katherine E. Tansey
MRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University, Cardiff, UK
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Alexander Teumer
Institute for Community Medicine, University Medicine Greifswald, Greifswald, Germany
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Anbupalam Thalamuthu
Centre for Healthy Brain Ageing, University of New South Wales, Sydney, Australia
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Sophia I. Thomopoulos
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Diana Tordesillas-Gutiérrez
Neuroimaging Unit, Technological Facilities, Valdecilla Biomedical Research Institute IDIVAL, Santander, SpainCentro Investigacion Biomedica en Red Salud Mental, Santander, Spain
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Jessica A. Turner
Department of Psychology, Georgia State University, Atlanta, USAMind Research Network, Albuquerque, USA
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Anne Uhlmann
Department of Psychiatry and Mental Health, University of Cape Town, Cape Town, South AfricaDepartment of Psychiatry, University of Vermont, Burlington, USA
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Costanza Ludovica Vallerga
Institute for Molecular Bioscience, The University of Queensland, Brisbane, Australia
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Dennis van der Meer
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, Norway
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Marjolein M.J. van Donkelaar
Department of Human Genetics, Radboud university medical center, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The Netherlands
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Liza van Eijk
School of Psychology, University of Queensland, Brisbane, AustraliaQueensland Brain Institute, University of Queensland, St Lucia, Australia
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Theo G.M. van Erp
Department of Psychiatry and Human Behavior, School of Medicine University of California, Irvine, Irvine, USA
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Neeltje E.M. van Haren
Department of Psychiatry, Brain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The NetherlandsDepartment of Child and Adolescent Psychiatry/Psychology, Erasmus Medical Center-Sophia Children’s Hospital, Rotterdam, The Netherlands
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Daan van Rooij
Department of Cognitive Neuroscience, Radboud university medical center, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The Netherlands
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Marie-José van Tol
Cognitive Neuroscience Center, Department of Neuroscience, University Medical Center Groningen, Groningen, The Netherlands
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Jan H. Veldink
Department of Neurology, Brain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The Netherlands
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Ellen Verhoef
Language and Genetics Department, Max Planck Institute for Psycholinguistics, Nijmegen, The Netherlands
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Esther Walton
Department of Psychology, Georgia State University, Atlanta, USAMRC Integrative Epidemiology Unit, Department of Population Health Sciences, Bristol Medical School, Bristol, UK
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Mingyuan Wang
Research Division, Institute of Mental Health, Singapore, Singapore
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Yunpeng Wang
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, Norway
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Joanna M. Wardlaw
Centre for Clinical Brain Sciences and Edinburgh Imaging, University of Edinburgh, Edinburgh, UKCentre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, UKUK Dementia Research Institute, The University of Edinburgh, Edinburgh, UK
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Wei Wen
Centre for Healthy Brain Ageing, University of New South Wales, Sydney, Australia
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Lars T. Westlye
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, NorwayDepartment of Psychology, University of Oslo, Oslo, Norway
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Christopher D. Whelan
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USADepartment of Molecular and Cellular Therapeutics, Royal College of Surgeons in Ireland, Dublin, Ireland
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Stephanie H. Witt
Department of Genetic Epidemiology in Psychiatry, Central Institute of Mental Health, Medical Faculty Mannheim, Heidelberg University, Mannheim, Germany
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Katharina Wittfeld
German Center for Neurodegenerative Diseases Rostock/Greifswald, Greifswald, GermanyDepartment of Psychiatry and Psychotherapy, University Medicine Greifswald, Greifswald, Germany
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Christiane Wolf
Department of Psychiatry, Psychosomatics and Psychotherapy, University of Würzburg, Würzburg, Germany
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Thomas Wolfers
Department of Human Genetics, Radboud university medical center, Nijmegen, The Netherlands
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Clarissa L. Yasuda
Department of Neurology, FCM, UNICAMP, Campinas, BrazilBRAINN - Brazilian Institute of Neuroscience and Neurotechnology, Campinas, Brazil
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Dario Zaremba
Department of Psychiatry, University of Münster, Münster, Germany
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Zuo Zhang
Social, Genetic and Developmental Psychiatry Centre, Institute of Psychiatry, Psychology & Neuroscience, King’s College London, London, UK
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Alyssa H. Zhu
Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Marcel P. Zwiers
Department of Cognitive Neuroscience, Radboud university medical center, Nijmegen, The NetherlandsDonders Centre for Cognitive Neuroimaging, Radboud University, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The Netherlands
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Eric Artiges
INSERM Unit 1000 - Neuroimaging & Psychiatry, Paris Saclay University,Gif sur Yvette, France
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Amelia A. Assareh
Centre for Healthy Brain Ageing, University of New South Wales, Sydney, Australia
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Rosa Ayesa-Arriola
Department of Psychiatry, University Hospital Marqués de Valdecilla, School of Medicine, University of Cantabria–IDIVAL, Santander, SpainCentro Investigacion Biomedica en Red Salud Mental, Santander, Spain
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Aysenil Belger
Duke UNC Brain Imaging and Analysis Center, Duke University Medical Center, Durham, USA
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Christine L. Brandt
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, Norway
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Gregory G. Brown
Department of Psychiatry, University of California San Diego, San Diego, USA
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Sven Cichon
Department of Biomedicine, University of Basel, Basel, SwitzerlandInstitute of Neuroscience and Medicine (INM-1), Research Centre Jülich, Jülich, GermanyInstitute of Medical Genetics and Pathology, University Hospital Basel, Basel, Switzerland
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Joanne E. Curran
Department of Human Genetics and South Texas Diabetes and Obesity Institute, University of Texas Rio Grande Valley School of Medicine, Brownsville, USA
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Gareth E. Davies
Avera Institute for Human Genetics, Sioux Falls, USA
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Franziska Degenhardt
Institute of Human Genetics, University of Bonn, School of Medicine & University Hospital Bonn, Bonn, Germany
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Bruno Dietsche
Department of Psychiatry and Psychotherapy, Philipps-University Marburg, Marburg, Germany
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Srdjan Djurovic
Department of Medical Genetics, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Department of Clinical Science, University of Bergen, Norway
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Colin P. Doherty
Department of Neurology, St James’s Hospital, Dublin, IrelandAcademic Unit of Neurology, TBSI, Dublin, IrelandFuture Neuro, Royal College of Surgeons in Ireland, Dublin, Ireland
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Ryan Espiritu
Information Sciences Institute, University of Southern California, Los Angeles, USA
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Daniel Garijo
Department of Neurology, St James’s Hospital, Dublin, Ireland
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Yolanda Gil
Information Sciences Institute, University of Southern California, Los Angeles, USA
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Penny A. Gowland
Sir Peter Mansfield Imaging Centre, University of Nottingham, Nottingham, UK
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Robert C. Green
Brigham and Women’s Hospital, Boston, USAThe Broad Institute, Boston, USAHarvard Medical School, Boston, USA
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Alexander N. Häusler
Center for Economics and Neuroscience, University of Bonn, Bonn, GermanyInstitute of Experimental Epileptology and Cognition Research, University Hospital Bonn, Germany
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Walter Heindel
Department of Clinical Radiology, University of Münster, Münster, Germany
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Beng-Choon Ho
Department of Psychiatry, University of Iowa College of Medicine, Iowa City, USA
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Wolfgang U. Hoffmann
Institute for Community Medicine, University Medicine Greifswald, Greifswald, GermanyGerman Center for Neurodegenerative Diseases Rostock/Greifswald, Greifswald, Germany
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Florian Holsboer
HMNC Holding GmbH, Munich, GermanyMax Planck Institute of Psychiatry, Munich, Germany
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Georg Homuth
University Medicine Greifswald, Interfaculty Institute for Genetics and Functional Genomics, Department of Functional Genomics, Greifswald, Germany
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Norbert Hosten
Institute of Diagnostic Radiology and Neuroradiology, Greifswald, Germany
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Clifford R. Jack
Dept of Radiology, Mayo Clinic, Rochester, USA
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MiHyun Jang
Information Sciences Institute, University of Southern California, Los Angeles, USA
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Andreas Jansen
Department of Psychiatry and Psychotherapy, Philipps-University Marburg, Marburg, GermanyCore-Unit Brainimaging, Faculty of Medicine, University of Marburg, Marburg, Germany
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Knut Kolskår
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, NorwayDepartment of Psychology, University of Oslo, Oslo, NorwaySunnaas Rehabilitation Hospital HT, Nesodden, Norway
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Sanne Koops
Department of Psychiatry, Brain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The Netherlands
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Axel Krug
Department of Psychiatry and Psychotherapy, Philipps-University Marburg, Marburg, Germany
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Kelvin O. Lim
Department of Psychiatry, University of Minnesota, Minneapolis, USA
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Jurjen J. Luykx
Department of Translational Neuroscience, Brain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The NetherlandsDepartment of Psychiatry, Brain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The NetherlandsDepartment of Psychiatry, ZNA hospitals, Antwerp, Belgium
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Daniel H. Mathalon
Department of Psychiatry and Weill Institute for Neurosciences, University of California San Francisco, San Francisco, USAMental Health Service 116d, Veterans Affairs San Francisco Healthcare System, San Francisco, USA
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Karen A. Mather
Centre for Healthy Brain Ageing, University of New South Wales, Sydney, AustraliaNeuroscience Research Australia, Sydney, Australia
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Venkata S. Mattay
Lieber Institute for Brain Development, Baltimore, USADepartment of Neurology, Johns Hopkins University, Baltimore, USADepartment of Radiology, Johns Hopkins University, Baltimore, USA
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Sarah Matthews
MRC Integrative Epidemiology Unit, Department of Population Health Sciences, Bristol Medical School, Bristol, UK
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Jaqueline Mayoral Van Son
Department of Psychiatry, University Hospital Marqués de Valdecilla, School of Medicine, University of Cantabria–IDIVAL, Santander, SpainCentro Investigacion Biomedica en Red Salud Mental, Santander, Spain
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Sarah C. McEwen
Department of Psychiatry, University of California San Diego, La Jolla, USAPacific Brain Health Center, Santa Monica, USAJohn Wayne Cancer Institute, Santa Monica, USA
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Ingrid Melle
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, Norway
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Derek W. Morris
Centre for Neuroimaging & Cognitive Genomics, National University of Ireland Galway, Galway, Ireland
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Bryon A. Mueller
Department of Psychiatry, University of Minnesota, Minneapolis, USA
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Matthias Nauck
Institute of Clinical Chemistry and Laboratory Medicine, University Medicine Greifswald, Greifswald, GermanyGerman Centre for Cardiovascular Research, Greifswald, Germany
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Jan E. Nordvik
Sunnaas Rehabilitation Hospital HT, Nesodden, Norway
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Markus M. Nöthen
Institute of Human Genetics, University of Bonn, School of Medicine & University Hospital Bonn, Bonn, Germany
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Daniel S. O’Leary
Department of Psychiatry, University of Iowa College of Medicine, Iowa City, USA
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Nils Opel
Department of Psychiatry, University of Münster, Münster, Germany
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Marie - Laure Paillère Martinot
INSERM Unit 1000 - Neuroimaging & Psychiatry, Paris Saclay University,Gif sur Yvette, FranceChild and adolescent psychiatry department, APHP Pitié Salpêtrière hospital, Paris, France
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G. Bruce Pike
Radiology and Clinical Neurosciences, Hotchkiss Brain Institute, University of Calgary, Calgary, Canada
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Adrian Preda
School of Medicine, University of California Irvine, Irvine, USA
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Erin B. Quinlan
Social, Genetic and Developmental Psychiatry Centre, Institute of Psychiatry, Psychology & Neuroscience, King’s College London, London, UK
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Varun Ratnakar
Information Sciences Institute, University of Southern California, Los Angeles, USA
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Simone Reppermund
Centre for Healthy Brain Ageing, University of New South Wales, Sydney, AustraliaDepartment of Developmental Disability Neuropsychiatry, University of New South Wales, Sydney, Australia
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Vidar M. Steen
NORMENT - K.G. Jebsen Centre for Psychosis Research, Department of Clinical Science, University of Bergen, NorwayDr. Einar Martens Research Group for Biological Psychiatry, Center for Medical Genetics and Molecular Medicine, Haukeland University Hospital, Bergen, Norway
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Fábio R. Torres
Department of Medical Genetics, School of Medical Sciences, University of Campinas - UNICAMP, Campinas, BrazilSection for Experimental Psychopathology and Neuroimaging, Department of General Psychiatry, Heidelberg University Hospital, Heidelberg, Germany
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Dick J. Veltman
Amsterdam UMC, Vrije Universiteit, Psychiatry, Amsterdam Public Health and Amsterdam Neuroscience and GGZinGeest, Amsterdam, The Netherlands
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James T. Voyvodic
Duke UNC Brain Imaging and Analysis Center, Duke University Medical Center, Durham, USA
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Robert Whelan
School of Psychology, Trinity College Dublin, Dublin, Ireland
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Tonya White
Department of Child and Adolescent Psychiatry/Psychology, Erasmus Medical Center-Sophia Children’s Hospital, Rotterdam, The NetherlandsDepartment of Radiology, Erasmus University Medical Centre, Rotterdam, The Netherlands
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Hidenaga Yamamori
Department of Psychiatry, Osaka University Graduate School of Medicine, Suita, Japan
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Oscar L. Lopez
Department of Epidemiology, Erasmus MC Medical Center, Rotterdam, The NetherlandsMedical Informatics, Erasmus MC Medical Center, Rotterdam, The Netherlands
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Hieab H.H. Adams
Department of Epidemiology, Erasmus MC Medical Center, Rotterdam, The NetherlandsDepartment of Radiology and Nuclear Medicine, Erasmus MC Medical Center, Rotterdam, The Netherlands
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Joshua C. Bis
Cardiovascular Health Research Unit, Department of Medicine, University of Washington, Seattle, USA
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Stephanie Debette
Inserm, Bordeaux Population Health Research Center, team VINTAGE, UMR 1219, University of Bordeaux, Bordeaux, FranceDepartment of Neurology, CHU de Bordeaux, Bordeaux, France
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Charles Decarli
Department of Neurology, University of California, Davis, Sacramento, USA
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Myriam Fornage
Institute of Molecular Medicine, University of Texas Health Science Center at Houston, Houston, USA
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Vilmundur Gudnason
Icelandic Heart Association, Kopavogur, IcelandFaculty of Medicine, University of Iceland, Reykjavik, Iceland
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Edith Hofer
Clinical Division of Neurogeriatrics, Department of Neurology, Medical University of Graz, Graz, AustriaInstitute for Medical Informatics, Statistics and Documentation, Medical University of Graz, Graz, Austria
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M. Arfan Ikram
Department of Epidemiology, Erasmus MC Medical Center, Rotterdam, The Netherlands
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Lenore Launer
Laboratory of Epidemiology and Population Sciences, Intramural Research Program, National Institute on Aging, Bethesda, USA
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W. T. Longstreth
Departments of Neurology and Epidemiology, University of Washington, Seattle, USA
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Bernard Mazoyer
Neurodegeneratives Diseases Institute UMR 5293, CNRS, CEA, University of Bordeaux, Bordeaux, France
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Thomas H. Mosley
MIND Center, University of Mississippi Medical Center, Jackson, USA
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Gennady V. Roshchupkin
Department of Epidemiology, Erasmus MC Medical Center, Rotterdam, The NetherlandsMedical Informatics, Erasmus MC Medical Center, Rotterdam, The NetherlandsDepartment of Radiology and Nuclear Medicine, Erasmus MC Medical Center, Rotterdam, The Netherlands
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Claudia L. Satizabal
Glenn Biggs Institute for Alzheimer’s and Neurodegenerative Diseases, University of Texas Health Sciences Center, San Antonio, USADepartment of Epidemiology & Biostatistics, University of Texas Health Sciences Center, San Antonio, USADepartment of Neurology, Boston University School of Medicine, Boston, USA
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Reinhold Schmidt
Departments of Genetics and Neuroscience, Yale University School of Medicine, New Haven, USA
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Sudha Seshadri
Glenn Biggs Institute for Alzheimer’s and Neurodegenerative Diseases, University of Texas Health Sciences Center, San Antonio, USADepartment of Neurology, Boston University School of Medicine, Boston, USAFramingham Heart Study and Department of Neurology, Boston University School of Medicine, Boston, USA
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Qiong Yang
Department of Biostatistics, Boston University School of Public Health, Boston, USA
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Marina K.M. Alvim
Department of Neurology, FCM, UNICAMP, Campinas, BrazilBRAINN - Brazilian Institute of Neuroscience and Neurotechnology, Campinas, Brazil
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David Ames
Academic Unit for Psychiatry of Old Age, University of Melbourne, Melbourne, AustraliaNational Ageing Research Institute, Melbourne, Australia
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Tim J. Anderson
Department of Medicine, University of Otago, Christchurch, Christchurch, New ZealandNew Zealand Brain Research Institute, Christchurch, New ZealandBrain Research New Zealand - Rangahau Roro Aotearoa, Christchurch, New ZealandDepartment of Neurology, Canterbury District Health Board, Christchurch, New Zealand
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Ole A. Andreassen
NORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, Norway
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Alejandro Arias-Vasquez
Department of Human Genetics, Radboud university medical center, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The NetherlandsDepartment of Psychiatry, Radboud university medical center, Nijmegen, The Netherlands
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Mark E. Bastin
Centre for Clinical Brain Sciences and Edinburgh Imaging, University of Edinburgh, Edinburgh, UKCentre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, UK
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Bernhard T. Baune
Department of Psychiatry, The University of Melbourne, Melbourne, Australia
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John Blangero
Department of Human Genetics and South Texas Diabetes and Obesity Institute, University of Texas Rio Grande Valley School of Medicine, Brownsville, USA
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Dorret I. Boomsma
Department of Biological Psychology, Vrije Universiteit Amsterdam, Amsterdam, The Netherlands
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Henry Brodaty
Centre for Healthy Brain Ageing, University of New South Wales, Sydney, AustraliaDementia Centre for Research Collaboration, University of New South Wales, Sydney, Australia
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Han G. Brunner
Department of Human Genetics, Radboud university medical center, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The NetherlandsDepartment of Clinical Genetics and School for Oncology & Developmental Biology (GROW), Maastricht University Medical Center, Maastricht, The Netherlands
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Randy L. Buckner
Department of Psychology and Center for Brain Science, Harvard University, Boston, USADepartment of Psychiatry, Massachusetts General Hospital, Boston, USADepartment of Radiology, Massachusetts General Hospital, Boston, USA
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Jan K. Buitelaar
Donders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The NetherlandsDepartment of Cognitive Neuroscience, Radboud university medical center, Nijmegen, The NetherlandsKarakter Child and Adolescent Psychiatry University Center, Nijmegen, The Netherlands
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Juan R. Bustillo
Department of Psychiatry, University of New Mexico, Albuquerque, USA
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Wiepke Cahn
Department of Psychiatry, University Medical Center Utrecht, Utrecht University, Utrecht, The Netherlands
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Vince Calhoun
Department of Electrical and Computer Engineering, The University of New Mexico, Albuquerque, USAMind Research Network, Albuquerque, USA
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Xavier Caseras
MRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University, Cardiff, UK
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Svenja Caspers
Institute of Neuroscience and Medicine (INM-1), Research Centre Jülich, Jülich, GermanyJARA-BRAIN, Jülich-Aachen Research Alliance, Jülich, GermanyInstitute for Anatomy I, Medical Faculty, Heinrich-Heine University, Düsseldorf, Germany
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Gianpiero L. Cavalleri
Molecular and Cellular Therapeutics, The Royal College of Surgeons In Ireland, Dublin, IrelandThe SFI FutureNeuro Research Centre, Dublin, Ireland
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Fernando Cendes
BRAINN - Brazilian Institute of Neuroscience and Neurotechnology, Campinas, BrazilDepartment of Neurology, FCM, UNICAMP, Campinas, Brazil
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Benedicto Crespo-Facorro
Centro Investigacion Biomedica en Red Salud Mental, Santander, SpainDonders Centre for Cognitive Neuroimaging, Radboud University, Nijmegen, The Netherlands
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John C. Dalrymple-Alford
New Zealand Brain Research Institute, Christchurch, New ZealandBrain Research New Zealand - Rangahau Roro Aotearoa, Christchurch, New ZealandDepartment of Psychology, University of Canterbury, Christchurch, New Zealand
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Udo Dannlowski
Department of Psychiatry, University of Münster, Münster, Germany
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Eco J.C. de Geus
Department of Biological Psychology, Vrije Universiteit Amsterdam, Amsterdam, The Netherlands
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Ian J. Deary
Department of Psychology, University of Edinburgh, Edinburgh, UKCentre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, UK
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Chantal Depondt
Department of Neurology, Hôpital Erasme, Université Libre de Bruxelles, Brussels, Belgium
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Sylvane Desrivières
Centre for Population Neuroscience and Precision Medicine (PONS), Institute of Psychiatry, Psychology and Neuroscience, King’s College London, London, UKSocial, Genetic and Developmental Psychiatry Centre, Institute of Psychiatry, Psychology & Neuroscience, King’s College London, London, UK
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Gary Donohoe
Centre for Neuroimaging & Cognitive Genomics, National University of Ireland Galway, Galway, Ireland
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Thomas Espeseth
Department of Psychology, University of Oslo, Oslo, NorwayNORMENT - K.G. Jebsen Centre for Psychosis Research, Division of Mental Health and Addiction, Oslo University Hospital, Oslo, Norway
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Guillén Fernández
Donders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The NetherlandsDepartment of Cognitive Neuroscience, Radboud university medical center, Nijmegen, The Netherlands
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Simon E. Fisher
Donders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The NetherlandsLanguage and Genetics Department, Max Planck Institute for Psycholinguistics, Nijmegen, The Netherlands
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Herta Flor
Department of Cognitive and Clinical Neuroscience, Central Institute of Mental Health, Medical Faculty Mannheim, Heidelberg University, Mannheim, Germany
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Andreas J. Forstner
Institute of Human Genetics, University of Bonn, School of Medicine & University Hospital Bonn, Bonn, GermanyDepartment of Biomedicine, University of Basel, Basel, SwitzerlandInstitute of Medical Genetics and Pathology, University Hospital Basel, Basel, SwitzerlandDepartment of Psychiatry (UPK), University of Basel, Basel, Switzerland
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Clyde Francks
Donders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The NetherlandsLanguage and Genetics Department, Max Planck Institute for Psycholinguistics, Nijmegen, The Netherlands
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Barbara Franke
Department of Human Genetics, Radboud university medical center, Nijmegen, The NetherlandsDonders Institute for Brain, Cognition and Behaviour, Radboud University, Nijmegen, The NetherlandsDepartment of Psychiatry, Radboud university medical center, Nijmegen, The Netherlands
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David C. Glahn
Department of Psychiatry, Yale University School of Medicine, New Haven, USAOlin Neuropsychiatric Research Center, Institute of Living, Hartford Hospital, Hartford, USA
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Randy L. Gollub
Department of Psychiatry, Harvard Medical School, Boston, USADepartment of Psychiatry, Massachusetts General Hospital, Boston, USADepartment of Radiology, Massachusetts General Hospital, Boston, USA
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Hans J. Grabe
Department of Psychiatry and Psychotherapy, University Medicine Greifswald, Greifswald, GermanyGerman Center for Neurodegenerative Diseases Rostock/Greifswald, Greifswald, Germany
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Oliver Gruber
Section for Experimental Psychopathology and Neuroimaging, Department of General Psychiatry, Heidelberg University Hospital, Heidelberg, Germany
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Asta K. Håberg
Department of Neuroscience, Norwegian University of Science and Technology, Trondheim, NorwayDepartment of Radiology and Nuclear medicine, Trondheim, Norway
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Ahmad R. Hariri
Department of Psychology and Neuroscience, Duke University, Durham, USA
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Catharina A. Hartman
Department of Psychiatry, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands
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Ryota Hashimoto
Department of Psychiatry, Osaka University Graduate School of Medicine, Suita, JapanMolecular Research Center for Children’s Mental Development, United Graduate School of Child Development, Osaka University, Suita, JapanDepartment of Pathology of Mental Diseases, National Institute of Mental Health, National Center of Neurology and Psychiatry, Tokyo, Japan
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Andreas Heinz
Department of Psychiatry and Psychotherapy, Campus Charité Mitte, Charité - Universitätsmedizin Berlin corporate member of Freie Universität Berlin, Humboldt-Universität zu Berlin, and Berlin Institute of Health, Berlin, Germany
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Manon H.J. Hillegers
Department of Child and Adolescent Psychiatry/Psychology, Erasmus Medical Center-Sophia Children’s Hospital, Rotterdam, The NetherlandsBrain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The Netherlands
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Pieter J. Hoekstra
Department of Psychiatry, University Medical Center Groningen, University of Groningen, Groningen, The Netherlands
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Avram J. Holmes
Department of Psychiatry, Massachusetts General Hospital, Boston, USADepartment of Psychology, Yale University, New Haven, USA
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L. Elliot Hong
Maryland Psychiatry Research Center, Department of Psychiatry, University of Maryland School of Medicine, Baltimore, USA
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William D. Hopkins
Neuroscience Institute, Georgia State University, Atlanta, USADivision of Developmental and Cognitive Neuroscience, Yerkes National Primate Research Center, Atlanta, USA
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Hilleke E. Hulshoff Pol
Department of Psychiatry, Brain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The Netherlands
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Terry L. Jernigan
Department of Radiology, University of California San Diego, San Diego, USADepartment of Cognitive Science, University of California San Diego, San Diego, USADepartment of Psychiatry, University of California San Diego, San Diego, USACenter for Human Development, University of California San Diego, La Jolla, USA
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Erik G. Jönsson
Centre for Psychiatric Research, Department of Clinical Neuroscience, Karolinska Institutet, Stockholm, SwedenNORMENT - K.G. Jebsen Centre for Psychosis Research, Institute of Clinical Medicine, University of Oslo, Oslo, Norway
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René S. Kahn
Department of Psychiatry, Icahn School of Medicine at Mount Sinai, New York, USADepartment of Psychiatry, Brain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The Netherlands
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Martin A. Kennedy
Department of Pathology and Biomedical Science, University of Otago, Christchurch, Christchurch, New Zealand
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Tilo T.J. Kircher
Department of Psychiatry and Psychotherapy, Philipps-University Marburg, Marburg, Germany
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Peter Kochunov
Maryland Psychiatry Research Center, Department of Psychiatry, University of Maryland School of Medicine, Baltimore, USA
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John B.J. Kwok
Neuroscience Research Australia, Sydney, AustraliaSchool of Medical Sciences, University of New South Wales, Sydney, AustraliaNeurogenetics and Epigenetics, Brain and Mind Centre, The University of Sydney, Sydney, Australia
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Stephanie Le Hellard
NORMENT - K.G. Jebsen Centre for Psychosis Research, Department of Clinical Science, University of Bergen, NorwayDr. Einar Martens Research Group for Biological Psychiatry, Center for Medical Genetics and Molecular Medicine, Haukeland University Hospital, Bergen, Norway
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INSERM Unit 1000 - Neuroimaging & Psychiatry, Paris Saclay University,Gif sur Yvette, France
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Centre for Neuroimaging & Cognitive Genomics, National University of Ireland Galway, Galway, Ireland
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Herston Imaging Research Facility, School of Clinical Sciences, Queensland University of Technology, Brisbane, Australia
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Department of Psychiatry and Psychotherapy, Central Institute of Mental Health Mannheim, Medical Faculty Mannheim, Heidelberg University, Mannheim, Germany
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Lars Nyberg
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Emma Children’s Hospital Academic Medical Center, Amsterdam, The NetherlandsDepartment of Pediatrics, Vrije Universiteit Medical Center, Vrije Universiteit Amsterdam, Amsterdam, The NetherlandsClinical Neuropsychology section, Vrije Universiteit Amsterdam, Amsterdam, The Netherlands
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Bloorview Research Institute, University of Toronto, Toronto, CanadaDepartments of Psychology and Psychiatry, University of Toronto, Toronto, CanadaCentre for Developing Brain, Child Mind Institute, New York City, USA
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Amsterdam UMC, Vrije Universiteit, Psychiatry, Amsterdam Public Health and Amsterdam Neuroscience and GGZinGeest, Amsterdam, The Netherlands
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Max Planck Institute of Psychiatry, Munich, Germany
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Centre for Population Neuroscience and Precision Medicine (PONS), Institute of Psychiatry, Psychology and Neuroscience, King’s College London, London, UKSocial, Genetic and Developmental Psychiatry Centre, Institute of Psychiatry, Psychology & Neuroscience, King’s College London, London, UK
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General Psychiatry, Institute of Mental Health, Singapore, Singapore
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Psychiatric and Neurodevelopmental Genetics Unit, Center for Genomic Medicine, Massachusetts General Hospital, Boston, USADepartment of Psychiatry, Massachusetts General Hospital, Boston, USAStanley Center for Psychiatric Research, Broad Institute, Boston, USA
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Cognitive Neuroscience Center, Department of Neuroscience, University Medical Center Groningen, Groningen, The NetherlandsDepartment of Psychiatry, University Medical Center Groningen, University of Groningen, Groningen, The NetherlandsBrain Center Rudolf Magnus, University Medical Center Utrecht, Utrecht University, Utrecht, The NetherlandsDepartment of Medical and Biological Psychology, University of Bergen, Bergen, Norway
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Department of Biological Psychology, Vrije Universiteit Amsterdam, Amsterdam, The Netherlands
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Institute for Community Medicine, University Medicine Greifswald, Greifswald, Germany
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Division of Mind and Brain Research, Department of Psychiatry and Psychotherapy, Campus Charité Mitte, Charité - Universitätsmedizin Berlin corporate member of Freie Universität Berlin, Humboldt-Universität zu Berlin, and Berlin Institute of Health, Berlin, Germany
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Queensland Brain Institute, University of Queensland, St Lucia, AustraliaCentre for Advanced Imaging, University of Queensland, Brisbane, Australia
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Center for Cognitive Neuroscience, Neuroscience and behavioral disorders program, Duke-National University of Singapore Medical School, Singapore, Singapore
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Department of Genetics & UNC Neuroscience Center, University of North Carolina at Chapel Hill, Chapel Hill, USA
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Imaging Genetics Center, Mark and Mary Stevens Neuroimaging and Informatics Institute, Keck School of Medicine of USC, University of Southern California, Los Angeles, USA
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Abstract

The cerebral cortex underlies our complex cognitive capabilities, yet we know little about the specific genetic loci influencing human cortical structure. To identify genetic variants impacting cortical structure, we conducted a genome-wide association meta-analysis of brain MRI data from 35,660 individuals with replication in 15,578 individuals. We analysed the surface area and average thickness of the whole cortex and 34 regions with known functional specialisations. We identified 206 nominally significant loci (P ≤ 5 × 10−8); 150 survived multiple testing correction (P ≤ 8.3 × 10−10; 140 surface area; 10 thickness). We found significant enrichment for loci influencing total surface area within regulatory elements active during prenatal cortical development, supporting the radial unit hypothesis. Loci impacting regional surface area cluster near genes in Wnt signalling pathways, known to influence progenitor expansion and areal identity. Variation in cortical structure is genetically correlated with cognitive function, Parkinson’s disease, insomnia, depression and ADHD.

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The genetic architecture of the human cerebral cortex
Katrina L. Grasby, Neda Jahanshad, Jodie N. Painter, Lucía Colodro-Conde, Janita Bralten, Derrek P. Hibar, Penelope A. Lind, Fabrizio Pizzagalli, Christopher R.K. Ching, Mary Agnes B. McMahon, Natalia Shatokhina, Leo C.P. Zsembik, Ingrid Agartz, Saud Alhusaini, Marcio A.A. Almeida, Dag Alnæs, Inge K. Amlien, Micael Andersson, Tyler Ard, Nicola J. Armstrong, Allison Ashley-Koch, Manon Bernard, Rachel M. Brouwer, Elizabeth E.L. Buimer, Robin Bülow, Christian Bürger, Dara M. Cannon, Mallar Chakravarty, Qiang Chen, Joshua W. Cheung, Baptiste Couvy-Duchesne, Anders M. Dale, Shareefa Dalvie, Tânia K. de Araujo, Greig I. de Zubicaray, Sonja M.C. de Zwarte, Anouk den Braber, Nhat Trung Doan, Katharina Dohm, Stefan Ehrlich, Hannah-Ruth Engelbrecht, Susanne Erk, Chun Chieh Fan, Iryna O. Fedko, Sonya F. Foley, Judith M. Ford, Masaki Fukunaga, Melanie E. Garrett, Tian Ge, Sudheer Giddaluru, Aaron L. Goldman, Nynke A. Groenewold, Dominik Grotegerd, Tiril P. Gurholt, Boris A. Gutman, Narelle K. Hansell, Mathew A. Harris, Marc B. Harrison, Courtney C. Haswell, Michael Hauser, Stefan Herms, Dirk J. Heslenfeld, New Fei Ho, David Hoehn, Per Hoffmann, Laurena Holleran, Martine Hoogman, Jouke-Jan Hottenga, Masashi Ikeda, Deborah Janowitz, Iris E. Jansen, Tianye Jia, Christiane Jockwitz, Ryota Kanai, Sherif Karama, Dalia Kasperaviciute, Tobias Kaufmann, Sinead Kelly, Masataka Kikuchi, Marieke Klein, Michael Knapp, Annchen R. Knodt, Bernd Krämer, Max Lam, Thomas M. Lancaster, Phil H. Lee, Tristram A. Lett, Lindsay B. Lewis, Iscia Lopes-Cendes, Michelle Luciano, Fabio Macciardi, Andre F. Marquand, Samuel R. Mathias, Tracy R. Melzer, Yuri Milaneschi, Nazanin Mirza-Schreiber, Jose C.V. Moreira, Thomas W. Mühleisen, Bertram Müller-Myhsok, Pablo Najt, Soichiro Nakahara, Kwangsik Nho, Loes M. Olde Loohuis, Dimitri Papadopoulos Orfanos, John F. Pearson, Toni L. Pitcher, Benno Pütz, Anjanibhargavi Ragothaman, Faisal M. Rashid, Ronny Redlich, Céline S. Reinbold, Jonathan Repple, Geneviève Richard, Brandalyn C. Riedel, Shannon L. Risacher, Cristiane S. Rocha, Nina Roth Mota, Lauren Salminen, Arvin Saremi, Andrew J. Saykin, Fenja Schlag, Lianne Schmaal, Peter R. Schofield, Rodrigo Secolin, Chin Yang Shapland, Li Shen, Jean Shin, Elena Shumskaya, Ida E. Sønderby, Emma Sprooten, Lachlan T. Strike, Katherine E. Tansey, Alexander Teumer, Anbupalam Thalamuthu, Sophia I. Thomopoulos, Diana Tordesillas-Gutiérrez, Jessica A. Turner, Anne Uhlmann, Costanza Ludovica Vallerga, Dennis van der Meer, Marjolein M.J. van Donkelaar, Liza van Eijk, Theo G.M. van Erp, Neeltje E.M. van Haren, Daan van Rooij, Marie-José van Tol, Jan H. Veldink, Ellen Verhoef, Esther Walton, Mingyuan Wang, Yunpeng Wang, Joanna M. Wardlaw, Wei Wen, Lars T. Westlye, Christopher D. Whelan, Stephanie H. Witt, Katharina Wittfeld, Christiane Wolf, Thomas Wolfers, Clarissa L. Yasuda, Dario Zaremba, Zuo Zhang, Alyssa H. Zhu, Marcel P. Zwiers, Eric Artiges, Amelia A. Assareh, Rosa Ayesa-Arriola, Aysenil Belger, Christine L. Brandt, Gregory G. Brown, Sven Cichon, Joanne E. Curran, Gareth E. Davies, Franziska Degenhardt, Bruno Dietsche, Srdjan Djurovic, Colin P. Doherty, Ryan Espiritu, Daniel Garijo, Yolanda Gil, Penny A. Gowland, Robert C. Green, Alexander N. Häusler, Walter Heindel, Beng-Choon Ho, Wolfgang U. Hoffmann, Florian Holsboer, Georg Homuth, Norbert Hosten, Clifford R. Jack Jr., MiHyun Jang, Andreas Jansen, Knut Kolskår, Sanne Koops, Axel Krug, Kelvin O. Lim, Jurjen J. Luykx, Daniel H. Mathalon, Karen A. Mather, Venkata S. Mattay, Sarah Matthews, Jaqueline Mayoral Van Son, Sarah C. McEwen, Ingrid Melle, Derek W. Morris, Bryon A. Mueller, Matthias Nauck, Jan E. Nordvik, Markus M. Nöthen, Daniel S. O’Leary, Nils Opel, Marie - Laure Paillère Martinot, G. Bruce Pike, Adrian Preda, Erin B. Quinlan, Varun Ratnakar, Simone Reppermund, Vidar M. Steen, Fábio R. Torres, Dick J. Veltman, James T. Voyvodic, Robert Whelan, Tonya White, Hidenaga Yamamori, Oscar L. Lopez, Hieab H.H. Adams, Joshua C. Bis, Stephanie Debette, Charles Decarli, Myriam Fornage, Vilmundur Gudnason, Edith Hofer, M. Arfan Ikram, Lenore Launer, W. T. Longstreth, Bernard Mazoyer, Thomas H. Mosley, Gennady V. Roshchupkin, Claudia L. Satizabal, Reinhold Schmidt, Sudha Seshadri, Qiong Yang, The Alzheimer’s Disease Neuroimaging Initiative, CHARGE Consortium, EPIGEN consortium, IMAGEN consortium, SYS consortium, Parkinson’s Progression Markers Initiative (PPMI), Marina K.M. Alvim, David Ames, Tim J. Anderson, Ole A. Andreassen, Alejandro Arias-Vasquez, Mark E. Bastin, Bernhard T. Baune, John Blangero, Dorret I. Boomsma, Henry Brodaty, Han G. Brunner, Randy L. Buckner, Jan K. Buitelaar, Juan R. Bustillo, Wiepke Cahn, Vince Calhoun, Xavier Caseras, Svenja Caspers, Gianpiero L. Cavalleri, Fernando Cendes, Benedicto Crespo-Facorro, John C. Dalrymple-Alford, Udo Dannlowski, Eco J.C. de Geus, Ian J. Deary, Chantal Depondt, Sylvane Desrivières, Gary Donohoe, Thomas Espeseth, Guillén Fernández, Simon E. Fisher, Herta Flor, Andreas J. Forstner, Clyde Francks, Barbara Franke, David C. Glahn, Randy L. Gollub, Hans J. Grabe, Oliver Gruber, Asta K. Håberg, Ahmad R. Hariri, Catharina A. Hartman, Ryota Hashimoto, Andreas Heinz, Manon H.J. Hillegers, Pieter J. Hoekstra, Avram J. Holmes, L. Elliot Hong, William D. Hopkins, Hilleke E. Hulshoff Pol, Terry L. Jernigan, Erik G. Jönsson, René S. Kahn, Martin A. Kennedy, Tilo T.J. Kircher, Peter Kochunov, John B.J. Kwok, Stephanie Le Hellard, Nicholas G. Martin, Jean-Luc Martinot, Colm McDonald, Katie L. McMahon, Andreas Meyer-Lindenberg, Rajendra A. Morey, Lars Nyberg, Jaap Oosterlaan, Roel A. Ophoff, Tomas Paus, Zdenka Pausova, Brenda W.J.H. Penninx, Tinca J.C. Polderman, Danielle Posthuma, Marcella Rietschel, Joshua L. Roffman, Laura M. Rowland, Perminder S. Sachdev, Philipp G. Sämann, Gunter Schumann, Kang Sim, Sanjay M. Sisodiya, Jordan W. Smoller, Iris E. Sommer, Beate St Pourcain, Dan J. Stein, Arthur W. Toga, Julian N. Trollor, Nic J.A. Van der Wee, Dennis van’t Ent, Henry Völzke, Henrik Walter, Bernd Weber, Daniel R. Weinberger, Margaret J. Wright, Juan Zhou, Jason L. Stein, Paul M. Thompson, Sarah E. Medland, on behalf of the Enhancing NeuroImaging Genetics through Meta-Analysis Consortium - Genetics working group
bioRxiv 399402; doi: https://doi.org/10.1101/399402
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The genetic architecture of the human cerebral cortex
Katrina L. Grasby, Neda Jahanshad, Jodie N. Painter, Lucía Colodro-Conde, Janita Bralten, Derrek P. Hibar, Penelope A. Lind, Fabrizio Pizzagalli, Christopher R.K. Ching, Mary Agnes B. McMahon, Natalia Shatokhina, Leo C.P. Zsembik, Ingrid Agartz, Saud Alhusaini, Marcio A.A. Almeida, Dag Alnæs, Inge K. Amlien, Micael Andersson, Tyler Ard, Nicola J. Armstrong, Allison Ashley-Koch, Manon Bernard, Rachel M. Brouwer, Elizabeth E.L. Buimer, Robin Bülow, Christian Bürger, Dara M. Cannon, Mallar Chakravarty, Qiang Chen, Joshua W. Cheung, Baptiste Couvy-Duchesne, Anders M. Dale, Shareefa Dalvie, Tânia K. de Araujo, Greig I. de Zubicaray, Sonja M.C. de Zwarte, Anouk den Braber, Nhat Trung Doan, Katharina Dohm, Stefan Ehrlich, Hannah-Ruth Engelbrecht, Susanne Erk, Chun Chieh Fan, Iryna O. Fedko, Sonya F. Foley, Judith M. Ford, Masaki Fukunaga, Melanie E. Garrett, Tian Ge, Sudheer Giddaluru, Aaron L. Goldman, Nynke A. Groenewold, Dominik Grotegerd, Tiril P. Gurholt, Boris A. Gutman, Narelle K. Hansell, Mathew A. Harris, Marc B. Harrison, Courtney C. Haswell, Michael Hauser, Stefan Herms, Dirk J. Heslenfeld, New Fei Ho, David Hoehn, Per Hoffmann, Laurena Holleran, Martine Hoogman, Jouke-Jan Hottenga, Masashi Ikeda, Deborah Janowitz, Iris E. Jansen, Tianye Jia, Christiane Jockwitz, Ryota Kanai, Sherif Karama, Dalia Kasperaviciute, Tobias Kaufmann, Sinead Kelly, Masataka Kikuchi, Marieke Klein, Michael Knapp, Annchen R. Knodt, Bernd Krämer, Max Lam, Thomas M. Lancaster, Phil H. Lee, Tristram A. Lett, Lindsay B. Lewis, Iscia Lopes-Cendes, Michelle Luciano, Fabio Macciardi, Andre F. Marquand, Samuel R. Mathias, Tracy R. Melzer, Yuri Milaneschi, Nazanin Mirza-Schreiber, Jose C.V. Moreira, Thomas W. Mühleisen, Bertram Müller-Myhsok, Pablo Najt, Soichiro Nakahara, Kwangsik Nho, Loes M. Olde Loohuis, Dimitri Papadopoulos Orfanos, John F. Pearson, Toni L. Pitcher, Benno Pütz, Anjanibhargavi Ragothaman, Faisal M. Rashid, Ronny Redlich, Céline S. Reinbold, Jonathan Repple, Geneviève Richard, Brandalyn C. Riedel, Shannon L. Risacher, Cristiane S. Rocha, Nina Roth Mota, Lauren Salminen, Arvin Saremi, Andrew J. Saykin, Fenja Schlag, Lianne Schmaal, Peter R. Schofield, Rodrigo Secolin, Chin Yang Shapland, Li Shen, Jean Shin, Elena Shumskaya, Ida E. Sønderby, Emma Sprooten, Lachlan T. Strike, Katherine E. Tansey, Alexander Teumer, Anbupalam Thalamuthu, Sophia I. Thomopoulos, Diana Tordesillas-Gutiérrez, Jessica A. Turner, Anne Uhlmann, Costanza Ludovica Vallerga, Dennis van der Meer, Marjolein M.J. van Donkelaar, Liza van Eijk, Theo G.M. van Erp, Neeltje E.M. van Haren, Daan van Rooij, Marie-José van Tol, Jan H. Veldink, Ellen Verhoef, Esther Walton, Mingyuan Wang, Yunpeng Wang, Joanna M. Wardlaw, Wei Wen, Lars T. Westlye, Christopher D. Whelan, Stephanie H. Witt, Katharina Wittfeld, Christiane Wolf, Thomas Wolfers, Clarissa L. Yasuda, Dario Zaremba, Zuo Zhang, Alyssa H. Zhu, Marcel P. Zwiers, Eric Artiges, Amelia A. Assareh, Rosa Ayesa-Arriola, Aysenil Belger, Christine L. Brandt, Gregory G. Brown, Sven Cichon, Joanne E. Curran, Gareth E. Davies, Franziska Degenhardt, Bruno Dietsche, Srdjan Djurovic, Colin P. Doherty, Ryan Espiritu, Daniel Garijo, Yolanda Gil, Penny A. Gowland, Robert C. Green, Alexander N. Häusler, Walter Heindel, Beng-Choon Ho, Wolfgang U. Hoffmann, Florian Holsboer, Georg Homuth, Norbert Hosten, Clifford R. Jack Jr., MiHyun Jang, Andreas Jansen, Knut Kolskår, Sanne Koops, Axel Krug, Kelvin O. Lim, Jurjen J. Luykx, Daniel H. Mathalon, Karen A. Mather, Venkata S. Mattay, Sarah Matthews, Jaqueline Mayoral Van Son, Sarah C. McEwen, Ingrid Melle, Derek W. Morris, Bryon A. Mueller, Matthias Nauck, Jan E. Nordvik, Markus M. Nöthen, Daniel S. O’Leary, Nils Opel, Marie - Laure Paillère Martinot, G. Bruce Pike, Adrian Preda, Erin B. Quinlan, Varun Ratnakar, Simone Reppermund, Vidar M. Steen, Fábio R. Torres, Dick J. Veltman, James T. Voyvodic, Robert Whelan, Tonya White, Hidenaga Yamamori, Oscar L. Lopez, Hieab H.H. Adams, Joshua C. Bis, Stephanie Debette, Charles Decarli, Myriam Fornage, Vilmundur Gudnason, Edith Hofer, M. Arfan Ikram, Lenore Launer, W. T. Longstreth, Bernard Mazoyer, Thomas H. Mosley, Gennady V. Roshchupkin, Claudia L. Satizabal, Reinhold Schmidt, Sudha Seshadri, Qiong Yang, The Alzheimer’s Disease Neuroimaging Initiative, CHARGE Consortium, EPIGEN consortium, IMAGEN consortium, SYS consortium, Parkinson’s Progression Markers Initiative (PPMI), Marina K.M. Alvim, David Ames, Tim J. Anderson, Ole A. Andreassen, Alejandro Arias-Vasquez, Mark E. Bastin, Bernhard T. Baune, John Blangero, Dorret I. Boomsma, Henry Brodaty, Han G. Brunner, Randy L. Buckner, Jan K. Buitelaar, Juan R. Bustillo, Wiepke Cahn, Vince Calhoun, Xavier Caseras, Svenja Caspers, Gianpiero L. Cavalleri, Fernando Cendes, Benedicto Crespo-Facorro, John C. Dalrymple-Alford, Udo Dannlowski, Eco J.C. de Geus, Ian J. Deary, Chantal Depondt, Sylvane Desrivières, Gary Donohoe, Thomas Espeseth, Guillén Fernández, Simon E. Fisher, Herta Flor, Andreas J. Forstner, Clyde Francks, Barbara Franke, David C. Glahn, Randy L. Gollub, Hans J. Grabe, Oliver Gruber, Asta K. Håberg, Ahmad R. Hariri, Catharina A. Hartman, Ryota Hashimoto, Andreas Heinz, Manon H.J. Hillegers, Pieter J. Hoekstra, Avram J. Holmes, L. Elliot Hong, William D. Hopkins, Hilleke E. Hulshoff Pol, Terry L. Jernigan, Erik G. Jönsson, René S. Kahn, Martin A. Kennedy, Tilo T.J. Kircher, Peter Kochunov, John B.J. Kwok, Stephanie Le Hellard, Nicholas G. Martin, Jean-Luc Martinot, Colm McDonald, Katie L. McMahon, Andreas Meyer-Lindenberg, Rajendra A. Morey, Lars Nyberg, Jaap Oosterlaan, Roel A. Ophoff, Tomas Paus, Zdenka Pausova, Brenda W.J.H. Penninx, Tinca J.C. Polderman, Danielle Posthuma, Marcella Rietschel, Joshua L. Roffman, Laura M. Rowland, Perminder S. Sachdev, Philipp G. Sämann, Gunter Schumann, Kang Sim, Sanjay M. Sisodiya, Jordan W. Smoller, Iris E. Sommer, Beate St Pourcain, Dan J. Stein, Arthur W. Toga, Julian N. Trollor, Nic J.A. Van der Wee, Dennis van’t Ent, Henry Völzke, Henrik Walter, Bernd Weber, Daniel R. Weinberger, Margaret J. Wright, Juan Zhou, Jason L. Stein, Paul M. Thompson, Sarah E. Medland, on behalf of the Enhancing NeuroImaging Genetics through Meta-Analysis Consortium - Genetics working group
bioRxiv 399402; doi: https://doi.org/10.1101/399402

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