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- September 29, 2018.
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- 1Institute of Evolutionary Biology, University of Edinburgh, Charlotte Auerbach Road, Edinburgh, United Kingdom
- 2Centre for Infection, Evolution and Immunity, University of Edinburgh, Edinburgh, United Kingdom
- ↵*Author for correspondence Email DC: danang.crysnanto{at}usys.ethz.ch DJO: darren.obbard{at}ed.ac.uk