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- May 20, 2019.
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- 1Laboratory for Bioinformatics Research, RIKEN Center for Biosystems Dynamics Research, Japan
- 2The Hakubi Center for Advanced Research/Department of Applied Mathematics and Physics, Graduate School of Informatics, Kyoto University, Japan
- 3Department of Biotechnology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Japan
- 4Bioinformatics Course, Master’s/Doctoral Program in Life Science Innovation (T-LSI), School of Integrative and Global Majors (SIGMA), University of Tsukuba, Japan
- ↵* Correspondence: itoshi.nikaido{at}riken.jp
† Equal contributor