1Solomon H. Snyder Department of Neuroscience, Johns Hopkins University School of Medicine, Baltimore, MD, 21205, USA
2Department of Ophthalmology, Johns Hopkins University School of Medicine, Baltimore, MD, 21205, USA
3Department of Neurology, Johns Hopkins University School of Medicine, Baltimore, MD, 21205, USA
4Center for Human Systems Biology, Johns Hopkins University School of Medicine, Baltimore, MD, 21205, USA
6Kavli Neuroscience Discovery Institute, Johns Hopkins University School of Medicine, Baltimore, MD, 21205, USA