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- August 11, 2019.
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- 1Plant Biology Graduate Program, Huck Institutes of the Life Sciences, The Pennsylvania State University, University Park, PA 16802, USA
- 2Leibniz-Institut DSMZ - Deutsche Sammlung von Mikroorganismen und Zellkulturen GmbH, Plant Virus Department, 38124 Braunschweig, Germany
- 3Department of Biology, The Pennsylvania State University, University Park, PA 16802, USA
- 4Istituto per la Protezione Sostenibile delle Piante, CNR, Strada delle Cacce 73, 10135, Torino, Italy
- 5Department of Plant Pathology and Environmental Microbiology, The Pennsylvania State University, University Park, PA 16802, USA
- ↵*Corresponding author: czr2{at}psu.edu