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- August 21, 2019.
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- 1Institute of Evolutionary Biology, University of Edinburgh, Charlotte Auerbach Road, Edinburgh, United Kingdom
- 2Charles Perkins Center, The University of Sydney, NSW, Australia
- 3Biosciences, College of Life & Environmental Sciences, University of Exeter, Penryn Campus, Penryn, Cornwall, United Kingdom
- 4Department of Evolutionary Biology & Systematic Zoology, Institute of Biochemistry and Biology, University of Potsdam, 4476 Potsdam, Germany
- ↵*Author for correspondence darren.obbard{at}ed.ac.uk
↵+ Contributed equally