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- October 8, 2019.
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- 1Marine Biology, Department of Biology, Ghent University, Krijgslaan 281 – S8, 9000 Gent, Belgium
- 2Cologne Biocenter, University of Cologne, Zülpicher Straβe 47b, 50674 Köln, Germany
- 3Department Aquatic Ecosystem Analysis, Helmholtz Centre for Environmental Research, Brückstraβe 3a, 39118 Magdeburg, Germany
- 4Department River Ecology, Helmholtz Centre for Environmental Research, Brückstraβe 3a, 39118 Magdeburg, Germany
- 5Flanders Marine Institute (VLIZ), Wandelaarkaai 7, 8400 Oostende, Belgium
- ↵*Author for correspondence (jens.boyen{at}ugent.be).