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- May 13, 2017.
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- Version 1 (June 15, 2016 - 10:50).
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The copyright holder for this preprint is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under a CC-BY 4.0 International license.
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- aDepartment of Zoology, Radboud University, Huygensgebouw, Heyendaalseweg 135. 6525 AJ Nijmegen, The Netherlands
- bDepartment of Zoology, University of Heidelberg, Im Neuenheimer Feld 234 D-69120 Heidelberg, Germany
- cDepartment of Biological Sciences, Purdue University, 915 W. State Street, West Lafayette, IN 47907, USA