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- October 31, 2017.
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- 1Department of Biology, 208 Erwin W. Mueller Laboratories, The Pennsylvania State University, University Park, PA 16802.
- 2Current address: 606 Fairchild Center, Department of Biological Sciences, Columbia University, New York, NY 1002.
- 3Department of Biology, University of Utah, Salt Lake City, UT 84112.
- 4Current address: Department of Genetics, University of Wisconsin, Madison, WI 53706.
- ↵**address correspondence to: Nitin Phadnis, 257 South 1400 East, Department of Biology, University of Utah, Salt Lake City, UT 84112; tel: (801)585-0493 email: nitin.phadnis{at}utah.edu
↵* these authors contributed equally