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- July 19, 2018.
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The copyright holder for this preprint is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under a CC-BY 4.0 International license.
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- 1Department of Botany, University of British Columbia, Vancouver, British Columbia V6T 1Z4, Canada
- 2Biodiversity Research Centre and Botany Department, University of British Columbia, Vancouver, British Columbia V6T 1Z4, Canada
- 3Departments of Botany and Zoology, University of British Columbia, Vancouver, British Columbia V6T 1Z4, Canada
- ↵4Author for correspondence (mgbontrager{at}gmail.com)