1 Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, Canada V6T 1Z4; Departments of Zoology and Biodiversity Research Centre, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, Canada V6T 1Z4 and Department of Biological Sciences, Macquarie University, Sydney NSW 2109, Australia wayne.maddison@ubc.ca.
2 Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, Canada V6T 1Z4; Departments of Zoology and Biodiversity Research Centre, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, Canada V6T 1Z4 and Department of Biological Sciences, Macquarie University, Sydney NSW 2109, Australia Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, Canada V6T 1Z4; Departments of Zoology and Biodiversity Research Centre, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, Canada V6T 1Z4 and Department of Biological Sciences, Macquarie University, Sydney NSW 2109, Australia.