Article Information
History
- November 8, 2017.
Copyright
The copyright holder for this preprint is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under a CC-BY-NC-ND 4.0 International license.
Author Information
- 1Nuffield Department of Medicine, University of Oxford, Oxford, United Kingdom
- 2Department of Ecology and Evolutionary Biology, Princeton University, Princeton, USA
- 3Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom
↵‡ These authors contributed equally to this work.